Darwinism: An Exposition of the Theory of Natural Selection — Alfred Russel Wallace
You may copy it, give it away or re-use it under the terms of the Project Gutenberg License included with this eBook or online at www.gutenberg.org . If you are not located in the United States, you will have to check the laws of the country where you are located before using this eBook. Title : Darwinism (1889) Author : Alfred Russel Wallace Release date : January 2, 2005 [eBook #14558] Most recently updated: October 28, 2024 Language : English Other information and formats : www.gutenberg.org/ebooks/14558 Credits : Produced by StevenGibbs and the Online Distributed Proofreading Team *** START OF D., F.L.S., ETC. WITH A PORTRAIT OF THE AUTHOR, MAP AND ILLUSTRATIONS MACMILLAN AND CO. LONDON AND NEW YORK 1889 Alfred R. Wallace PREFACE TO SECOND EDITION The present edition is a reprint of the first, with a few verbal corrections and the alteration of some erroneous or doubtful statements. Of these latter the following are the most important:— P. 30 . The statement as to the fulmar petrel, which Professor A. Newton assures me is erroneous, has been modified. P. 34 . A note is added as to Darwin's statement about the missel and song-thrushes in Scotland. P. 172 . An error as to the differently-coloured herds of cattle in the Falkland Islands, is corrected. PARKSTONE, DORSET August, 1889 . PREFACE TO FIRST EDITION The present work treats the problem of the Origin of Species on the same general lines as were adopted by Darwin; but from the standpoint reached after nearly thirty years of discussion, with an abundance of new facts and the advocacy of many new or old theories. While not attempting to deal, even in outline, with the vast subject of evolution in general, an endeavour has been made to give such an account of the theory of Natural Selection as may enable any intelligent reader to obtain a clear conception of Darwin's work, and to understand something of the power and range of his great principle. Darwin wrote for a generation which had not accepted evolution, and which poured contempt on those who upheld the derivation of species from species by any natural law of descent. He did his work so well that "descent with modification" is now universally accepted as the order of nature in the organic world; and the rising generation of naturalists can hardly realise the novelty of this idea, or that their fathers considered it a scientific heresy to be condemned rather than seriously discussed. The objections now made to Darwin's theory apply, solely, to the particular means by which the change of species has been brought about, not to the fact of that change. The objectors seek to minimise the agency of natural selection and to subordinate it to laws of variation, of use and disuse, of intelligence, and of heredity. These views and objections are urged with much force and more confidence, and for the most part by the modern school of laboratory naturalists, to whom the peculiarities and distinctions of species, as such, their distribution and their affinities, have little interest as compared with the problems of histology and embryology, of physiology and morphology. Their work in these departments is of the greatest interest and of the highest importance, but it is not the kind of work which, by itself, enables one to form a sound judgment on the questions involved in the action of the law of natural selection. These rest mainly on the external and vital relations of species to species in a state of nature—on what has been well termed by Semper the "physiology of organisms," rather than on the anatomy or physiology of organs. It has always been considered a weakness in Darwin's work that he based his theory, primarily, on the evidence of variation in domesticated animals and cultivated plants. I have endeavoured to secure a firm foundation for the theory in the variations of organisms in a state of nature; and as the exact amount and precise character of these variations is of paramount importance in the numerous problems that arise when we apply the theory to explain the facts of nature, I have endeavoured, by means of a series of diagrams, to exhibit to the eye the actual variations as they are found to exist in a sufficient number of species. By doing this, not only does the reader obtain a better and more precise idea of variation than can be given by any number of tabular statements or cases of extreme individual variation, but we obtain a basis of fact by which to test the statements and objections usually put forth on the subject of specific variability; and it will be found that, throughout the work, I have frequently to appeal to these diagrams and the facts they illustrate, just as Darwin was accustomed to appeal to the facts of variation among dogs and pigeons. I have also made what appears to me an important change in the arrangement of the subject. Instead of treating first the comparatively difficult and unfamiliar details of variation, I commence with the Struggle for Existence, which is really the fundamental phenomenon on which natural selection depends, while the particular facts which illustrate it are comparatively familiar and very interesting. It has the further advantage that, after discussing variation and the effects of artificial selection, we proceed at once to explain how natural selection acts. Among the subjects of novelty or interest discussed in this volume, and which have important bearings on the theory of natural selection, are: (1) A proof that all specific characters are (or once have been) either useful in themselves or correlated with useful characters (Chap. VI); (2) a proof that natural selection can, in certain cases, increase the sterility of crosses (Chap. VII); (3) a fuller discussion of the colour relations of animals, with additional facts and arguments on the origin of sexual differences of colour (Chaps. VIII-X); (4) an attempted solution of the difficulty presented by the occurrence of both very simple and very complex modes of securing t